Do inter and intra require hyphens?

No, hyphens are typically not required when using the prefixes “inter” and “intra.”

However, some style guides advise using hyphens in certain instances. For example, the AP Style Guide suggests using a hyphen when the root word is a proper noun (e.g., “inter-American”), and the Chicago Manual of Style recommends including a hyphen to avoid a double “a” or “i” (e.g., “intra-arterial”).